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- Coupling-control ratio + ('''Coupling-control ratios''' ''CCR'' are … '''Coupling-control ratios''' ''CCR'' are [[flux control ratio]]s ''FCR'' at a constant mitochondrial [[pathway-control state]]. In mitochondrial preparations, there are three well-defined coupling states of respiration: [[LEAK respiration]], [[OXPHOS]], and [[Electron transfer pathway |Electron-transfer-pathway state]] (ET state). In these states, the corresponding respirtory rates are symbolized as ''L'', ''P'', and ''E''. In living cells, the OXPHOS state cannot be induced, but in the [[ROUTINE]] state the respiration rate is ''R''. A reference rate ''Z'' is defined by taking ''Z'' as the maximum flux, i.e. flux ''E'' in the ET-state, such that the lower and upper limits of the ''CCR'' are defined as 0.0 and 1.0. Then there are two mitochondrial ''CCR'', [[L/E |''L/E'']] and [[P/E |''P/E'']], and two ''CCR'' for living cells, [[L/E |''L/E'']] and [[ROUTINE-control ratio |''R/E'']].[[ROUTINE-control ratio |''R/E'']].)